March 28, 2018

Let’s Talk Lighting

by John Tullock

Roughly 30 years ago, I wrote a series of articles for Freshwater and Marine Aquarium magazine entitled “Light In the Marine Aquarium.” The article had been prompted by the then-new idea of reef tanks, and the limited selection of appropriate lighting systems. The problem was simply that hobbyists and the industry had not recognized the importance of light to a functioning ecosystem.

After that article appeared, a long-running debate ensued regarding the “best” lighting system for a reef tank. The reef tank craze completely eclipsed freshwater aquarium keeping for a while, and all lighting technology was marketed toward that group of hobbyists. Today, with freshwater planted aquariums enjoying a resurgence in popularity, the lighting debate has been re-kindled, albeit with a different perspective. With that in mind, let’s take a look at what matters and what doesn’t in your choice of an aquarium lighting system.

All photosynthetic organisms, whether they are Cryptocorynes or the symbiotic zooxanthellae that live in the tissues of hermatypic corals, depend upon chlorophyll to harness light energy for the manufacture of sugar molecules. Chlorophyll occurs in several molecular configurations, with the most universal one being chlorophyll a. All photosynthetic organisms contain chlorophyll a. The second most common form of chlorophyll is found in plants and algae, while the less common forms are found in different groups of micro-organisms, including cyanobacteria. Certain other pigments play a minor role in energy capture. We will confine our discussion here to chlorophylls a and b.



Pictured above is a graphic representation of energy absorption by chlorophylls a and b. Note that most of the absorption is in the blue and red regions of the spectrum, with four distinct peaks, two for each type of chlorophyll.

 

The second image presents the spectral output of a typical T5 fluorescent lamp. The details are not important, but note that not all of the light from this lamp is at the wavelengths absorbed by chlorophyll. This is the case with any type of artificial lighting, only a portion of the available light energy is at the wavelengths plants need.

This opens up the question of how best to choose appropriate artificial lighting for a freshwater planted tank, or indeed any situation where plants are to grow without sunshine. The answer lies in a physical parameter known as “photosynthetically active radiation,” or PAR value. PAR value is reported as micromoles of photons per square meter per second,” and represents our best direct measurement of light output from any putative plant-growth light source.

Let’s be clear: PAR is the ONLY direct measure of a given light source’s suitability for plants. Other terms associated with lighting are often tossed about in this context. Here are some of them:

Watts per gallon—this is an outdated method that has obvious flaws and has been abandoned by serious aquarists.

Kelvin temperature—a measurement of the color-rendering ability of a light source, as compared to natural daylight, that says nothing about the ability of the light to support plant growth. As a general rule, higher Kelvin temperature light sources emit a high proportion of their light in the blue wavelengths plants use, but they may also be deficient in the red wavelengths that are equally essential to plant growth and development.

Lux—a measurement of light intensity that says nothing about the spectral quality of the light.

PAR value can be easily determined by use of a PAR meter. Once prohibitively expensive, these instruments are now priced within the range of most hobbyist clubs and certainly most aquarium shops. In practice, a PAR reading of about 50 units at the level of the substrate directly beneath the center of the light source will support the growth of a wide range of popular aquarium plants. All PAR readings for aquarium purposes should be made with the sensor underwater.

Manufacturers have capitalized on the lighting debate of the last three decades, and many now put spectral data, Kelvin temperature ratings and so forth on their packaging. My advice: look for the ones with PAR values on the package, or better yet, measure those PAR values yourself and then decide.

LIGHT INTENSITY

“Using very strong lighting is like driving a fast car. It could get you to your destination more quickly, or end in an accident. Strong lighting coupled with organic waste or damaged plants trigger algae spores to bloom and the end result is visible algae in the tank.” Jeff Miotke

This is a guide to the appropriate light intensity as measured on the surface of the substrate
* 30umols of PAR (photosynthetic active radiation): Low light plants such as ferns, mosses, Cryptocoryne, Anubias, Bucephalandra species grow well. If you are growing purely shade plants, it is smart to stay in this range.
* 50umols of PAR will enable you to grow any commercially available aquatic plant (when coupled with optimal CO2 levels). However, this gives little allowance for shading issues in crowded tanks. Most nature style and Iwagumi style tanks that do not require high density do well at this range. 
* 50 to 150umols of PAR; Colored stems will display better coloration and carpets spread very fast as light levels increase, at the risk of more algae issues. Tank cleanliness and plant health must be maintained at optimal conditions to prevent run-away algae. Allows for increased planting density for light demanding plants. Most experienced aquarists can do well in this range with regular upkeep.
* 200umols of PAR & above; Marginal gains to plant growth form and coloration starts to be outweighed by large potential instability of algae outbreaks. Might be useful inducing stronger coloration in shorter stems close to substrate; and for farming purposes. Should only be used by aquarists used to maintaining already algae-free tanks.

Organism
Genus
Species
Lighting Requirment
PAR (µE m2sec)
Minimum
Ideal
Maximum
Plant
Anubias
angustifolia
 
40
125
200
Plant
Anubias
barteri
 
40
75
125
Plant
Anubias
nana
 
40
75
125
Plant
Anubis Species
 
40
100
200
Plant
Aponogeton Species
High
100
150
200
Plant
Bacopa
Monnieri
 
100
175
250
Plant
Bacopa Species
 
100
175
250
Plant
Cabomba
 caroliniana
Medium
100
150
200
Plant
Cabomba Species
High
100
150
250
Plant
Crinum Species
High
100
150
200
Plant
Cryptocoryne 
beckettii
 
50
125
200
Plant
Cryptocoryne 
wendtii
 
50
125
200
Plant
Cryptocorynes Species
Low
40
50
200
Plant
Echinodorous
barthii
High
150
200
250
Plant
Echinodorous
bleheri
Low – Very High
40
175
250
Plant
Echinodorous
macrophyllus
Average to High
50
150
200
Plant
Echinodorous
osiris
Average to High
50
150
200
Plant
Echinodorous
Ozelot
 
50
150
200
Plant
Echinodorous
palaefolius
 
100
200
250
Plant
Echinodorous
quadricostatus
 
50
150
200
Plant
Echinodorous
Rose
 
50
100
150
Plant
Echinodorous
Rubin
 
50
150
200
Plant
Echinodorous
tenellus
 
100
150
200
Plant
Echinodorous
uruguayensis
 
75
175
250
Plant
Echinodorous Species
Medium
50
100
150
Plant
Hygrophila
polysperma
 
150
200
250
Plant
Hygrophila Species
 
150
200
250
Plant
Lilaeopsis Species
High
100
150
200
Plant
Limnophila Species
High
100
150
200
Plant
Ludwigia Species
High
100
150
200
Plant
Microsorum
pteropus
 
40
125
200
Plant
Microsorum Species
 
40
125
200
Plant
Myriophyllum Species
High
100
150
200
Plant
Nesaea Species
High
100
150
200
Plant
Nymphaea Species 
High
100
150
200
Plant
Pogostemon Species
High
100
150
200
Plant
Rotala Species
High
100
150
200
Plant
Sagittaria Species
High
100
150
200
Plant
Versicularia
dubyana
 
40
125
200
Plant
Versicularia Species
 
40
125
200
Plant
Vallisneria 
americana
 
50
150
200
Plant
Vallisneria 
spiralis
 
40
150
200
Plant
Versicularia Species
 
40
150
200
Plant
Egeria
densa
 
100
150
250
Plant
Egeria Species
 
100
150
250
This list was produced in association with Arcadia products –  the experts in aquarium lighting.